For other uses, see Naja (disambiguation).
Indian cobra, N. naja
Naja, Laurenti, 1768
Naja is a genus of venomous elapid snakes. Although several other genera share the common name, Naja species are the most recognized and most widespread group of snakes commonly known as cobras. The genus Naja consists of 20 to 22 species, but has undergone several taxonomic revisions in recent years, so sources vary greatly. They range throughout Africa, Southwest Asia, South Asia and Southeast Asia. The most recent revison. listed 28 species after the synonymisation of Boulengerina and Paranaja with Naja
4.1 King cobra,
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The origin of this genus name is from the Old Indic nāga, cognate with English 'snake', Germanic: *snēk-a-, Proto-IE: *(s)nēg-o-.
Different Naja species vary in length and most are relatively slender-bodied snakes. Most species are capable of attaining lengths of 1.84 m (6.0 ft). Maximum lengths for some of the larger species of cobra are around 3.1 m (10 ft), with the forest cobra arguably being the longest species. All have a characteristic ability to raise the front quarters of their bodies off the ground and flatten their necks to appear larger to a potential predator.
All species in the genus Naja are capable of delivering a fatal bite to a human. Most species have strongly neurotoxic venom, which attacks the nervous system, causing paralysis, but many also have cytotoxic features which causes swelling and necrosis, and has a significant anticoagulant effect. Some also have cardiotoxic components to their venom.
Several Naja species, referred to as spitting cobras, have a specialized venom delivery mechanism, in which their front fangs, instead of releasing venom through the tips (similar to a hypodermic needle), have a rifled opening in the front surface which allows the snake to propel the venom out of the mouth. While typically referred to as "spitting", the action is more like squirting. The range and accuracy with which they can shoot their venom varies from species to species, but it is used primarily as a defense mechanism. Once sprayed onto a victim's skin, the venom acts as a severe irritant. If it is introduced to the eye, it can cause a severe burning sensation and temporary or even permanent blindness if not cleaned out immediately and thoroughly.
Murine subcutaneous LD50 values for some cobra species include 0.20 mg/kg for N. philippinensis (Philippine cobra), which is considered to be the most venomous species, 0.29--0.53 mg/kg for N. atra (Chinese cobra), 0.40 mg/kg for N. oxiana (Caspian cobra), 0.45--0.80 mg/kg for N. naja (Indian cobra), 0.72 mg/kg for N. nivea (Cape cobra), 1.15 mg/kg for N. haje (Egyptian cobra) and 2.0 mg/kg for Naja nigricollis (black-necked spitting cobra).
Some murine IV LD50 values include 0.29 mg/kg for N. melanoleuca (forest cobra), 0.345 mg/kg for N. atra (Chinese cobra), 0.373 mg/kg for N. kaouthia (monocled cobra), and 0.96 mg/kg for N. oxiana (Caspian cobra).
The murine IP LD50 of N. annulata (banded water cobra) and N. christyi (Congo water cobra) venoms were 0.143 and 0.120 mg/kg, respectively. Other IP LD50 values include N. haje (Egyptian cobra) at 0.185 mg/kg, N. kaouthia (monocled cobra) at 0.225 mg/kg, N. naja at 0.315 mg/kg, N. melanoleuca at 0.324 mg/kg, N. nivea (Cape cobra) at 0.4 mg/kg, N. nigricollis at 0.4 mg/kg, and N. pallida (red spitting cobra) at 2.0 mg/kg.
Top 10 venomous Naja species
Philippine cobra (N. philippinensis)
Caspian cobra (N. oxiana)
Forest cobra (N. melanoleuca)
Samar cobra (N. samarensis)
Monocled cobra (N. kaouthia)
Indochinese spitting cobra (N. siamensis)
Equatorial spitting cobra (N. sumatrana)
Chinese cobra (N. atra)
Cape cobra (N. nivea)
Indian cobra (N. naja)
Anchieta's cobra (Angolan Cobra)
Angola, Botswana, Namibia, Zambia, eastern Zimbabwe
(Buchholz and Peters, 1876)
Banded water cobra
Cameroon, Central African Republic, Democratic Republic of Congo, Republic of Congo, Equatorial Guinea, Gabon, Rwanda, and the province of Cabinda in Angola
Botswana, Malawi, Mozambique, South Africa, Swaziland, Zambia, Zimbabwe
Oman, Saudi Arabia, Yemen
Wüster and Broadley, 2007
Ashe's spitting cobra (giant spitting cobra)
Southern Ethiopia, Kenya, Somalia, eastern Uganda
Southern China, northern Laos, Taiwan, northern Vietnam
Congo water cobra
Democratic Republic of Congo, Republic of Congo, and in the province of Cabinda in Angola
Algeria, Burkina Faso, Cameroon, Central African Republic, northern Democratic Republic of the Congo, Chad, Egypt, Eritrea, Ethiopia, Kenya, Libya, Mali, Mauritania, Morocco, Niger, Nigeria, Somalia, Sudan, Tanzania, Togo, Tunisia, Uganda, Western Sahara
Bangladesh, Bhutan, Burma, Cambodia, southern China, eastern India, Laos, northwestern Malaysia, Nepal, Thailand, southeastern Tibet, Vietnam
Benin, Burkina Faso, Cameroon, Ghana, Guinea, Ivory Coast, Mali, Gambia, Mauritania, Niger, Nigeria, Senegal, Togo
Slowinski & Wüster, 2000
Mandalay spitting cobra (Burmese spitting cobra)
Angola, Benin, Burkina Faso, Burundi, Cameroon, Central African Republic, Chad, Democratic Republic of the Congo (Zaire), Congo, Ethiopia, Gabon, Ghana, Gambia, Guinea, Guinea Bissau, Ivory Coast, Kenya, Liberia, Malawi, Mali, Mozambique, Niger, Nigeria, Rwanda, Sao Tom`e, Senegal, Sierra Leone, Somalia, South Africa, Sudan, Tanzania, Togo, Uganda, Zambia, Zimbabwe
Mozambique spitting cobra
Extreme southeastern Angola, Botswana, Malawi, Mozambique, Somalia, northeastern Namibia, South Africa, Swaziland, Tanzania (including Pemba Island), Zambia, Zimbabwe
Cameroon, Congo, Democratic Republic of the Congo (Zaire), Gabon
Indian cobra (spectacled cobra)
Bangladesh, Bhutan, India, Nepal, Pakistan, Sri Lanka
Zebra spitting cobra
Angola, Namibia, South Africa
Black-necked spitting cobra
Angola, Benin, Burkina Faso, Burundi, Cameroon, Central African Republic, Chad, Democratic Republic of the Congo (except in the center), Congo, Ethiopia, Gabon, Gambia, Ghana, Guinea Bissau, Guinea, Ivory Coast, Kenya, Liberia, Mali, Mauritania, Namibia, Niger, Nigeria, Rwanda, Senegal, Sierra Leone, Sudan, Tanzania, Somalia, Togo, Uganda, Zambia
Cape cobra (yellow cobra)
Botswana, Lesotho, Namibia, South Africa
Wüster & Broadley, 2003
Nubian spitting cobra
Chad, Egypt, Eritrea, Niger, Sudan
Afghanistan, northwest India, Iran, Kyrgyzstan, Pakistan, Tajikistan, Turkmenistan, Uzbekistan
Red spitting cobra
Djibouti, Ethiopia, Kenya, Somalia, Tanzania
Philippines (Luzon, Mindoro)
India (Andaman Islands)
Philippines (Mindanao, Bohol, Leyte, Samar, Camiguin)
Trape, Chirio & Wüster, 2009
Benin, Burkina Faso, Ghana, Guinea, Mali, Niger, Nigeria, Senegal
Indo-Chinese spitting cobra
Cambodia, Laos, Thailand, Vietnam
F. Boie, 1827
Javan spitting cobra
Indonesia (Java, Lesser Sunda Islands, East Timor)
Equatorial spitting cobra
Brunei, Indonesia (Sumatra, Borneo, Bangka, Belitung), Malaysia, Philippines (Palawan), southern Thailand, Singapore
*) Not including the nominate subspecies., ) Type species.
Despite its popular name, the king cobra (Ophiophagus hannah) is not considered a member of the genus Naja.
The genus contains several species complexes of closely related and often similar species, some of them only recently described or defined. Several recent taxonomic studies have revealed species not included in the current listing in ITIS:
Naja anchietae (Bocage, 1879), or Anchieta's cobra, is regarded as a subspecies of N. haje by Mertens (1937) and of N. annulifera by Broadley (1995). It is regarded as a full species by Broadley and Wüster (2004).,
Naja arabica Scortecci, 1932, the Arabian cobra, has long been considered a subspecies of N. haje, but was recently raised to the status of species.,
Naja ashei Broadley and Wüster, 2007, Ashe's spitting cobra, is a newly described species found in Africa.,
Naja nigricincta Bogert, 1940, was long regarded as a subspecies of N. nigricollis, but was recently found to be a full species (with N. n. woodi as a subspecies).,
Naja senegalensis Trape et al., 2009, is a new species encompassing what were previously considered to be the West African savanna populations of N. haje.,
Two recent molecular phylogenetic studies have also supported the incorporation of the species normally assigned to the genera Boulengerina and Paranaja into Naja, as both are closely related to the forest cobra (Naja melanoleuca).
Wallach et al. suggested recognition of four subgenera within Naja: Naja for the Asiatic cobras, Boulengerina for the African forest, water and burrowing cobras, Uraeus for the Egyptian and Cape cobra group and Afronaja for the African spitting cobras.